植物学报 ›› 2018, Vol. 53 ›› Issue (4): 487-501.DOI: 10.11983/CBB17082
郭书磊1,2, 张君1, 齐建双1, 岳润清1, 韩小花1, 燕树锋1, 卢彩霞1, 傅晓雷1, 陈娜娜1, 库丽霞2,*(), 铁双贵1,*(
)
收稿日期:
2017-04-13
接受日期:
2017-10-25
出版日期:
2018-07-01
发布日期:
2018-09-11
通讯作者:
库丽霞,铁双贵
作者简介:
共同第一作者。
基金资助:
Guo Shulei1,2, Zhang Jun1, Qi Jianshuang1, Yue Runqing1, Han Xiaohua1, Yan Shufeng1, Lu Caixia1, Fu Xiaolei1, Chen Nana1, Ku Lixia2,*(), Tie Shuanggui1,*(
)
Received:
2017-04-13
Accepted:
2017-10-25
Online:
2018-07-01
Published:
2018-09-11
Contact:
Ku Lixia,Tie Shuanggui
About author:
These authors contributed equally to this paper
摘要:
叶长、叶宽、叶面积及叶夹角不仅影响玉米(Zea mays)光合效率, 也是株型的重要构成因素。通过对620个叶形QTL进行整合, 构建不同遗传背景下的叶形QTL整合图谱, 利用元分析发掘出22个叶长、22个叶宽、12个叶面积以及17个叶夹角mQTL; 进一步运用生物信息学手段, 确定44个与叶片发育密切相关的候选基因。分析发现, 仅有NAL7-like、YABBY6- like和GRF2等13个基因位于mQTL区间内, 而玉米中已克隆的KNOTTED1、AN3/GIF1、rgd1/lbl1、mwp1及SRL2-like、HYL1-like和CYCB2;4-like等水稻(Oryza sativa)和拟南芥(Arabidopsis thaliana)叶形同源基因位于未被整合的QTL内; 对44个候选基因在叶片长、宽、厚发育过程中基部-末端、中央-边缘、远轴-近轴的调控机理进行归纳分析, 发现玉米中仅有少数几个候选基因被报道, 揭示了叶形发育的部分分子机理。因此, 对玉米叶形相关mQTL/QTL及基因进行全面深入的分析, 不仅有助于增加对其遗传结构的了解, 发掘更多候选基因, 阐明叶形发育和形成的分子机制, 还可为耐密理想株型的分子标记辅助选择提供依据。
郭书磊, 张君, 齐建双, 岳润清, 韩小花, 燕树锋, 卢彩霞, 傅晓雷, 陈娜娜, 库丽霞, 铁双贵. 玉米叶形相关性状的Meta-QTL及候选基因分析. 植物学报, 2018, 53(4): 487-501.
Guo Shulei, Zhang Jun, Qi Jianshuang, Yue Runqing, Han Xiaohua, Yan Shufeng, Lu Caixia, Fu Xiaolei, Chen Nana, Ku Lixia, Tie Shuanggui. Analysis of Meta-quantitative Trait Loci and Their Candidate Genes Related to Leaf Shape in Maize. Chinese Bulletin of Botany, 2018, 53(4): 487-501.
Parents | Type of Pop. | Pop. size | No. of QTL for LL | No. of QTL for LW | No. of QTL for LAr | No. of QTL for LA | No./type of marker | Analysis method | Reference |
---|---|---|---|---|---|---|---|---|---|
B73×G79 | F2:3 | 214 | — | — | 7 | — | 185/RFLP | IM | Agrama et al., 1999 |
B73×Mo17 | RIL | 180 | — | — | — | 9 | 192/SSR | CIM | Mickelson et al., 2002 |
H21×Mo17 | F2:3 | 120 | — | — | — | 7 | 102/SSR | CIM | 于永涛等, 2006 |
Zi330×K36 | F2:3 | 114 | — | — | — | 2 | 90/SSR | CIM | 于永涛等, 2006 |
Ye478×Dan340 | F2:3 | 397 | — | — | — | 6 | 138/SSR | CIM | 路明等, 2007 |
Mo17×Huangzao4 | RIL | 239 | 4 | 2 | 5 | — | 98/SSR | CIM | 郑祖平等, 2007 |
Ye478×Wu312 | RIL | 218 | — | — | 7 | — | 184/SSR | ICIM | 刘建超等, 2010 |
Yu82×Shen137 | F2:3 | 229 | 3 | 4 | 13 | 3 | 222/SSR | CIM | Ku et al., 2010, 2012a |
Yu82×Yu87-1 | F2:3 | 256 | 5 | 5 | — | 5 | 216/SSR | CIM | Ku et al., 2012b |
B73/Mo17 etc. | NAM | 4892 | 36 | 34 | — | 30 | 203000/SNP | GWAS | Tian et al., 2011 |
N6×BT-1 | RIL | 250 | — | — | 6 | — | 207/SSR | CIM | 李贤唐等, 2011 |
Y105×Y106 | F2 | 189 | 5 | 7 | 6 | 8 | 215/SSR | ICIM | 郭莹, 2012 |
Y114×Y115 | F2 | 189 | 1 | 1 | 2 | 3 | 204/SSR | ICIM | 郭莹, 2012 |
Ye478×Wu312 | RIL | 218 | 14 | 7 | 9 | — | 184/SSR | CIM | Cai et al., 2012 |
Yu82×D132 | F2:3 | 245 | — | — | 18 | — | 204/SSR | CIM | Ku et al., 2012 |
B73×1212 | RIL | 325 | 67 | 62 | — | — | 208/SSR | ICIM | 唐登国, 2013 |
B73×Mo17 | RIL | 93 | 1 | 2 | — | 3 | IBM2 map | CIM | Wassom, 2013 |
CY5×YL106 | F2:3 | 144 | — | — | — | 8 | 212/SSR | CIM | 刘正等, 2014 |
Z58/87-1// PH6WC/Zi330 | CP | 228 | — | — | — | 13 | 225/SSR | IM | 张姿丽等, 2014b |
T4×T19 | F2:3 | 232 | — | — | 4 | — | 81/SSR | CIM | 张姿丽等, 2014a |
Yu82×Yu87-1 | RIL | 208 | — | 18 | — | — | 1370/SNP | CIM | Guo et al., 2015 |
Yu82×Shen137 | RIL | 197 | — | 9 | — | — | 1411/SNP | CIM | Guo et al., 2015 |
Zong3×Yu87-1 | RIL | 223 | — | 10 | — | — | 1479/SNP | CIM | Guo et al., 2015 |
Yu537A×Shen137 | RIL | 212 | — | 9 | — | — | 1371/SNP | CIM | Guo et al., 2015 |
B73×Mo17 | DH | 221 | 9 | 17 | 16 | 12 | 5935/bin markers | ICIM | 张志腾, 2015 |
Xu178×K12 | RIL | 150 | — | — | — | 34 | 191/SSR | CIM | 常立国等, 2016 |
M1-7×SYF | F2 | 259 | — | — | 36 | — | 218/SSR | CIM | 安允权等, 2016 |
Yu82×D132 | RIL | 234 | 5 | 7 | 4 | — | 1226/SNP | CIM | Wei et al., 2016 |
表1 玉米叶形相关QTL信息整合
Table 1 Summary of the QTL of leaf shape in maize reported previously
Parents | Type of Pop. | Pop. size | No. of QTL for LL | No. of QTL for LW | No. of QTL for LAr | No. of QTL for LA | No./type of marker | Analysis method | Reference |
---|---|---|---|---|---|---|---|---|---|
B73×G79 | F2:3 | 214 | — | — | 7 | — | 185/RFLP | IM | Agrama et al., 1999 |
B73×Mo17 | RIL | 180 | — | — | — | 9 | 192/SSR | CIM | Mickelson et al., 2002 |
H21×Mo17 | F2:3 | 120 | — | — | — | 7 | 102/SSR | CIM | 于永涛等, 2006 |
Zi330×K36 | F2:3 | 114 | — | — | — | 2 | 90/SSR | CIM | 于永涛等, 2006 |
Ye478×Dan340 | F2:3 | 397 | — | — | — | 6 | 138/SSR | CIM | 路明等, 2007 |
Mo17×Huangzao4 | RIL | 239 | 4 | 2 | 5 | — | 98/SSR | CIM | 郑祖平等, 2007 |
Ye478×Wu312 | RIL | 218 | — | — | 7 | — | 184/SSR | ICIM | 刘建超等, 2010 |
Yu82×Shen137 | F2:3 | 229 | 3 | 4 | 13 | 3 | 222/SSR | CIM | Ku et al., 2010, 2012a |
Yu82×Yu87-1 | F2:3 | 256 | 5 | 5 | — | 5 | 216/SSR | CIM | Ku et al., 2012b |
B73/Mo17 etc. | NAM | 4892 | 36 | 34 | — | 30 | 203000/SNP | GWAS | Tian et al., 2011 |
N6×BT-1 | RIL | 250 | — | — | 6 | — | 207/SSR | CIM | 李贤唐等, 2011 |
Y105×Y106 | F2 | 189 | 5 | 7 | 6 | 8 | 215/SSR | ICIM | 郭莹, 2012 |
Y114×Y115 | F2 | 189 | 1 | 1 | 2 | 3 | 204/SSR | ICIM | 郭莹, 2012 |
Ye478×Wu312 | RIL | 218 | 14 | 7 | 9 | — | 184/SSR | CIM | Cai et al., 2012 |
Yu82×D132 | F2:3 | 245 | — | — | 18 | — | 204/SSR | CIM | Ku et al., 2012 |
B73×1212 | RIL | 325 | 67 | 62 | — | — | 208/SSR | ICIM | 唐登国, 2013 |
B73×Mo17 | RIL | 93 | 1 | 2 | — | 3 | IBM2 map | CIM | Wassom, 2013 |
CY5×YL106 | F2:3 | 144 | — | — | — | 8 | 212/SSR | CIM | 刘正等, 2014 |
Z58/87-1// PH6WC/Zi330 | CP | 228 | — | — | — | 13 | 225/SSR | IM | 张姿丽等, 2014b |
T4×T19 | F2:3 | 232 | — | — | 4 | — | 81/SSR | CIM | 张姿丽等, 2014a |
Yu82×Yu87-1 | RIL | 208 | — | 18 | — | — | 1370/SNP | CIM | Guo et al., 2015 |
Yu82×Shen137 | RIL | 197 | — | 9 | — | — | 1411/SNP | CIM | Guo et al., 2015 |
Zong3×Yu87-1 | RIL | 223 | — | 10 | — | — | 1479/SNP | CIM | Guo et al., 2015 |
Yu537A×Shen137 | RIL | 212 | — | 9 | — | — | 1371/SNP | CIM | Guo et al., 2015 |
B73×Mo17 | DH | 221 | 9 | 17 | 16 | 12 | 5935/bin markers | ICIM | 张志腾, 2015 |
Xu178×K12 | RIL | 150 | — | — | — | 34 | 191/SSR | CIM | 常立国等, 2016 |
M1-7×SYF | F2 | 259 | — | — | 36 | — | 218/SSR | CIM | 安允权等, 2016 |
Yu82×D132 | RIL | 234 | 5 | 7 | 4 | — | 1226/SNP | CIM | Wei et al., 2016 |
Bin | mQTL/QTL | CI (cM) | Candidate gene | Annotation | Homologous gene | E-value | Reference |
---|---|---|---|---|---|---|---|
1.02 | mQTLW1-1 | 153.8-158.3 | GRMZM2G480386 | NAL7-like | Os03g0162000/YUCCA7 | 0 | Fujino et al., 2008 |
1.04 | q12SevLW1 | 270.6-286.2 | GRMZM2G011483 | SRL2-like | Os03g19520/SRL2 | 0 | Liu et al., 2016 |
1.05 | mQTLW1-2 | 485.9-505.6 | GRMZM2G141955 | YABBY6-like | Os12g42610/YAB6 | 3E-88 | Toriba et al., 2007; Zhang et al., 2009 |
GRMZM2G003509 | PHB-like | AT2G34710/AtHB14 | 0 | Kim et al., 2003; Mallory et al., 2004 | |||
1.09 | mQTLW1-3 | 820.6-845.5 | GRMZM2G018414 | GRF8 | AT4G37740/AtGRF2 | 4E-37 | Debernardi et al., 2014 |
1.10 | q4LWm139 | 873.7-902.1 | GRMZM2G017087 | KNOTTED1 | AT4G08150/KN1 | 2E-114 | Ramirez et al., 2009 |
GRMZM2G178261 | GRF1-like | AT2G22840/AtGRF1 | 2E-66 | Kim et al., 2003 | |||
GRMZM2G180246 | AN3/GIF1 | AT5G28640/AtAN3/GIF1 | 1E-36 | Nelissen et al., 2015 | |||
1.11 | q4LLm155 | 941.6-1022.3 | GRMZM2G135447 | OSH43-like | Os03g0771500/OsH43 | 4E-99 | Sentoku et al., 2000 |
2.02 | q3LAr2a | 67.5-79.7 | GRMZM2G174784 | AP2-like | AT4G36920/AP2 | 7E-102 | Würschum et al., 2006; Mlotshwa et al., 2006 |
2.02 | mQTLW2-1 | 118.3-147.8 | GRMZM2G102346 | NAL1-like | Os04g52479/NAL1 | 0 | Kubo et al., 2017 |
GRMZM2G041223 | GRF6 | AT3G13960/AtGRF5 | 1E-36 | Horiguchi et al., 2006; Debernardi et al., 2014 | |||
2.06 | q4RLAr2 | 355.7-374.4 | GRMZM2G444808 | HYL1-like | AT1G09700/HYL1 | 6E-56 | Liu et al., 2011 |
2.06 | q8SecLW2 | 370.0-381.6 | GRMZM2G083972 | LNG2-like | AT3G02170/LNG2 | 6E-57 | Lee et al., 2006 |
GRMZM2G161382 | CYCD3;3-like | AT3G50070/CYCD3;3 | 5E-56 | Dewitte et al., 2007 | |||
2.07 | q7LL2b | 450.0-461.9 | GRMZM2G004619 | GRF4 | AT4G37740/AtGRF2 | 5E-28 | Kim et al., 2003 |
2.10 | q12LAr2 | 713.1-732.4 | GRMZM2G146688 | ANT-like | AT4G37750/ANT | 1E-131 | Mizukami and Fis- cher, 2000 |
3.02 | q4LWm309 | 69.6-90.3 | GRMZM2G143235 | ROT3-like | AT4G36380/ROT3 | 1E-176 | Kim et al., 1998 |
3.06 | mQTLW3-2 | 368.5-390.3 | GRMZM2G118250 | AS2-like | AT1G65620/AS2 | 1E-66 | Iwakawa et al., 2007 |
3.08 | q4LWm410 | 613.6-653.7 | GRMZM2G437460 | ARF3-like | AT2G33860/ARF3/ETT | 1E-169 | Kelley et al., 2012 |
4.04 | q13NLW4 | 220.5-230.6 | GRMZM2G402653 | OSH1-like | Os03g0727000/OsH1 | 1E-75 | Matsuoka et al., 1993 |
4.06 | mQTLW4 | 373.7-399.4 | GRMZM2G124566 | GRF9-like | AT2G36400/AtGRF3 | 1E-36 | Kim et al., 2003 |
4.08 | q7LL4 | 429.5-451.0 | GRMZM2G075117 | CYCD3;1-like | AT4G34160/CYCD3;1 | 4E-47 | Dewitte et al., 2007 |
GRMZM2G105335 | GRF3 | AT3G13960/AtGRF5 | 8E-49 | Horiguchi et al., 2006; Debernardi et al., 2014 | |||
5.03 | q4LWm602 | 268.8-295.0 | GRMZM2G361518 | AGO10-like | AT5G43810/AGO10/ZLL | 0 | Zhu et al., 2011; Roodbarkelari et al., 2015 |
5.03 | q4LLm605 | 280.1-296.1 | GRMZM2G171349 | COW1-like | AT4G34580/COW1 | 3E-66 | Woo et al., 2007 |
5.06 | q4LLm661 | 500.7-521.1 | GRMZM2G034876 | GRF1 | AT3G13960/AtGRF5 | 1E-47 | Horiguchi et al., 2006; Debernardi et al., 2014 |
5.07 | mQTLW5-3 | 556.7-578.5 | GRMZM5G893117 | GRF9 | AT3G13960/AtGRF5 | 1E-27 | Horiguchi et al., 2006; Debernardi et al., 2014 |
6.01 | q8FirLW6-1 | 82.3-93.9 | GRMZM2G122537 | PRS1-like | AT2G28610/PRS1 | 6E-29 | Matsumoto and Ok- ada, 2001; Nakata et al., 2012 |
GRMZM2G020187 | rgd1/lbl1 | AT5G23570/SGS3 | 1E-157 | Dotto et al., 2014 | |||
mQTL/QTL | CI (cM) | Candidate gene | Annotation | Homologous gene | E-value | Reference | |
6.01 | mQTLAr6 | 92.8-108.9 | GRMZM2G098594 | GRF14 | AT3G13960/AtGRF5 | 1E-36 | Horiguchi et al., 2006; Debernardi et al., 2014 |
6.02 | q7PLL6 | 123.0-144.8 | GRMZM2G073671 | CYCB2;3-like | AT1G20610/CYCB2;3 | 1E-144 | Eloy et al., 2011 |
GRMZM2G157820 | CLF-like | AT2G23380/CLF | 0 | Menges et al., 2005; Schatlowski et al., 2010 | |||
6.04 | q4LWm706 | 158.2-205.1 | GRMZM5G850129 | GRF7 | AT3G13960/AtGRF5 | 4E-41 | Horiguchi et al., 2006; Debernardi et al., 2014 |
7.00 | q4LWm762 | 8.0-93.0 | GRMZM2G028041 | OSH15-like | Os07g0129700/OsH15 | 1.E-168 | Nagasaki et al., 2001 |
GRMZM2G019200 | DRL1-like | Os11g0312782/DRL1 | 0 | Jun et al., 2015 | |||
7.02 | q12EigLW7 | 200.7-217.9 | GRMZM2G107377 | CYCD3;2-like | AT5G67260/CYCD3;2 | 8E-49 | Dewitte et al., 2007 |
7.02 | q12FirLW7 | 229.3-246.3 | GRMZM2G082264 | mwp1 | Os09g23200/SLL1/Kan1 | 2E-170 | Candela et al., 2008 |
7.03 | q3LW7 | 380.6-498.2 | GRMZM2G096709 | GRF10 | AT4G37740/AtGRF2 | 4E-29 | Kim et al., 2003; Deb- ernardi et al., 2014 |
8.08 | q3LW8 | 599.1-603.8 | GRMZM5G874163 | ARF4-like | AT5G60450/ARF4 | 7E-120 | Pekker et al., 2005; Hunter et al., 2006 |
9.03 | mQTLW9 | 206.1-230.6 | GRMZM2G119359 | GRF12 | AT2G06200/AtGRF6 | 1E-30 | Kim et al., 2003; Liang et al., 2014 |
GRMZM5G870176 | NRL-like | Os12g36890/CSLD4 | 0 | Hu et al., 2010 | |||
10.04 | mQTLW10 | 295.6-327.3 | GRMZM2G078641 | GRF2 | GRAT3G13960/AtGRF7 | 3E-156 | Horiguchi et al., 2006; Debernardi et al., 2014 |
10.06 | q1HNLAr10 | 367.0-383.9 | GRMZM2G061287 | CYCB2;4-like | AT1G76310/CYCB2;4 | 9E-148 | Menges et al., 2005; Eloy et al., 2011 |
表2 mQTL/QTL区域内的玉米叶形候选基因
Table 2 Candidate genes for maize leaf shape of mQTL/QTL
Bin | mQTL/QTL | CI (cM) | Candidate gene | Annotation | Homologous gene | E-value | Reference |
---|---|---|---|---|---|---|---|
1.02 | mQTLW1-1 | 153.8-158.3 | GRMZM2G480386 | NAL7-like | Os03g0162000/YUCCA7 | 0 | Fujino et al., 2008 |
1.04 | q12SevLW1 | 270.6-286.2 | GRMZM2G011483 | SRL2-like | Os03g19520/SRL2 | 0 | Liu et al., 2016 |
1.05 | mQTLW1-2 | 485.9-505.6 | GRMZM2G141955 | YABBY6-like | Os12g42610/YAB6 | 3E-88 | Toriba et al., 2007; Zhang et al., 2009 |
GRMZM2G003509 | PHB-like | AT2G34710/AtHB14 | 0 | Kim et al., 2003; Mallory et al., 2004 | |||
1.09 | mQTLW1-3 | 820.6-845.5 | GRMZM2G018414 | GRF8 | AT4G37740/AtGRF2 | 4E-37 | Debernardi et al., 2014 |
1.10 | q4LWm139 | 873.7-902.1 | GRMZM2G017087 | KNOTTED1 | AT4G08150/KN1 | 2E-114 | Ramirez et al., 2009 |
GRMZM2G178261 | GRF1-like | AT2G22840/AtGRF1 | 2E-66 | Kim et al., 2003 | |||
GRMZM2G180246 | AN3/GIF1 | AT5G28640/AtAN3/GIF1 | 1E-36 | Nelissen et al., 2015 | |||
1.11 | q4LLm155 | 941.6-1022.3 | GRMZM2G135447 | OSH43-like | Os03g0771500/OsH43 | 4E-99 | Sentoku et al., 2000 |
2.02 | q3LAr2a | 67.5-79.7 | GRMZM2G174784 | AP2-like | AT4G36920/AP2 | 7E-102 | Würschum et al., 2006; Mlotshwa et al., 2006 |
2.02 | mQTLW2-1 | 118.3-147.8 | GRMZM2G102346 | NAL1-like | Os04g52479/NAL1 | 0 | Kubo et al., 2017 |
GRMZM2G041223 | GRF6 | AT3G13960/AtGRF5 | 1E-36 | Horiguchi et al., 2006; Debernardi et al., 2014 | |||
2.06 | q4RLAr2 | 355.7-374.4 | GRMZM2G444808 | HYL1-like | AT1G09700/HYL1 | 6E-56 | Liu et al., 2011 |
2.06 | q8SecLW2 | 370.0-381.6 | GRMZM2G083972 | LNG2-like | AT3G02170/LNG2 | 6E-57 | Lee et al., 2006 |
GRMZM2G161382 | CYCD3;3-like | AT3G50070/CYCD3;3 | 5E-56 | Dewitte et al., 2007 | |||
2.07 | q7LL2b | 450.0-461.9 | GRMZM2G004619 | GRF4 | AT4G37740/AtGRF2 | 5E-28 | Kim et al., 2003 |
2.10 | q12LAr2 | 713.1-732.4 | GRMZM2G146688 | ANT-like | AT4G37750/ANT | 1E-131 | Mizukami and Fis- cher, 2000 |
3.02 | q4LWm309 | 69.6-90.3 | GRMZM2G143235 | ROT3-like | AT4G36380/ROT3 | 1E-176 | Kim et al., 1998 |
3.06 | mQTLW3-2 | 368.5-390.3 | GRMZM2G118250 | AS2-like | AT1G65620/AS2 | 1E-66 | Iwakawa et al., 2007 |
3.08 | q4LWm410 | 613.6-653.7 | GRMZM2G437460 | ARF3-like | AT2G33860/ARF3/ETT | 1E-169 | Kelley et al., 2012 |
4.04 | q13NLW4 | 220.5-230.6 | GRMZM2G402653 | OSH1-like | Os03g0727000/OsH1 | 1E-75 | Matsuoka et al., 1993 |
4.06 | mQTLW4 | 373.7-399.4 | GRMZM2G124566 | GRF9-like | AT2G36400/AtGRF3 | 1E-36 | Kim et al., 2003 |
4.08 | q7LL4 | 429.5-451.0 | GRMZM2G075117 | CYCD3;1-like | AT4G34160/CYCD3;1 | 4E-47 | Dewitte et al., 2007 |
GRMZM2G105335 | GRF3 | AT3G13960/AtGRF5 | 8E-49 | Horiguchi et al., 2006; Debernardi et al., 2014 | |||
5.03 | q4LWm602 | 268.8-295.0 | GRMZM2G361518 | AGO10-like | AT5G43810/AGO10/ZLL | 0 | Zhu et al., 2011; Roodbarkelari et al., 2015 |
5.03 | q4LLm605 | 280.1-296.1 | GRMZM2G171349 | COW1-like | AT4G34580/COW1 | 3E-66 | Woo et al., 2007 |
5.06 | q4LLm661 | 500.7-521.1 | GRMZM2G034876 | GRF1 | AT3G13960/AtGRF5 | 1E-47 | Horiguchi et al., 2006; Debernardi et al., 2014 |
5.07 | mQTLW5-3 | 556.7-578.5 | GRMZM5G893117 | GRF9 | AT3G13960/AtGRF5 | 1E-27 | Horiguchi et al., 2006; Debernardi et al., 2014 |
6.01 | q8FirLW6-1 | 82.3-93.9 | GRMZM2G122537 | PRS1-like | AT2G28610/PRS1 | 6E-29 | Matsumoto and Ok- ada, 2001; Nakata et al., 2012 |
GRMZM2G020187 | rgd1/lbl1 | AT5G23570/SGS3 | 1E-157 | Dotto et al., 2014 | |||
mQTL/QTL | CI (cM) | Candidate gene | Annotation | Homologous gene | E-value | Reference | |
6.01 | mQTLAr6 | 92.8-108.9 | GRMZM2G098594 | GRF14 | AT3G13960/AtGRF5 | 1E-36 | Horiguchi et al., 2006; Debernardi et al., 2014 |
6.02 | q7PLL6 | 123.0-144.8 | GRMZM2G073671 | CYCB2;3-like | AT1G20610/CYCB2;3 | 1E-144 | Eloy et al., 2011 |
GRMZM2G157820 | CLF-like | AT2G23380/CLF | 0 | Menges et al., 2005; Schatlowski et al., 2010 | |||
6.04 | q4LWm706 | 158.2-205.1 | GRMZM5G850129 | GRF7 | AT3G13960/AtGRF5 | 4E-41 | Horiguchi et al., 2006; Debernardi et al., 2014 |
7.00 | q4LWm762 | 8.0-93.0 | GRMZM2G028041 | OSH15-like | Os07g0129700/OsH15 | 1.E-168 | Nagasaki et al., 2001 |
GRMZM2G019200 | DRL1-like | Os11g0312782/DRL1 | 0 | Jun et al., 2015 | |||
7.02 | q12EigLW7 | 200.7-217.9 | GRMZM2G107377 | CYCD3;2-like | AT5G67260/CYCD3;2 | 8E-49 | Dewitte et al., 2007 |
7.02 | q12FirLW7 | 229.3-246.3 | GRMZM2G082264 | mwp1 | Os09g23200/SLL1/Kan1 | 2E-170 | Candela et al., 2008 |
7.03 | q3LW7 | 380.6-498.2 | GRMZM2G096709 | GRF10 | AT4G37740/AtGRF2 | 4E-29 | Kim et al., 2003; Deb- ernardi et al., 2014 |
8.08 | q3LW8 | 599.1-603.8 | GRMZM5G874163 | ARF4-like | AT5G60450/ARF4 | 7E-120 | Pekker et al., 2005; Hunter et al., 2006 |
9.03 | mQTLW9 | 206.1-230.6 | GRMZM2G119359 | GRF12 | AT2G06200/AtGRF6 | 1E-30 | Kim et al., 2003; Liang et al., 2014 |
GRMZM5G870176 | NRL-like | Os12g36890/CSLD4 | 0 | Hu et al., 2010 | |||
10.04 | mQTLW10 | 295.6-327.3 | GRMZM2G078641 | GRF2 | GRAT3G13960/AtGRF7 | 3E-156 | Horiguchi et al., 2006; Debernardi et al., 2014 |
10.06 | q1HNLAr10 | 367.0-383.9 | GRMZM2G061287 | CYCB2;4-like | AT1G76310/CYCB2;4 | 9E-148 | Menges et al., 2005; Eloy et al., 2011 |
图1 整合图谱中叶形相关mQTL在玉米各染色体上的分布Chr: 染色体。染色体上红色区域为mQTL重叠部分所在染色体的位置; 染色体左侧为整合后mQTL的名称及位置分布; 染色体右侧为整合图谱中的标记及遗传图距(cM)。
Figure 1 Distribution of leaf shape mQTL on maize chromosomes in the integrated mapChr: Chromosome. The red region is the identified location of the overlaps of mQTL on the chromosome; The position distribution and the name for mQTL are listed in the left of chromosome; Marker name and genetic distance (cM) are listed in the right of chromosome.
图2 玉米44个叶形候选基因的氨基酸序列聚类进化树方框标注的为玉米中已克隆且功能已知的叶形候选基因蛋白功能
Figure 2 Phylogenetic trees of amino acid sequences of 44 candidate genes related to leaf shape of maizeFunction of candidate genes which have been known and cloned related to maize leaf shape are highlighted in the box
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