木灵藓科分类与系统学研究:历史、问题和展望
# 共同第一作者
收稿日期: 2016-03-12
录用日期: 2016-10-16
网络出版日期: 2017-04-05
基金资助
国家自然科学基金(No.31570208, No.31370233, No.30970184, No.30570121)
Taxonomic Revision and Phylogenetics of the Orthotrichaceae: History, Problems and Prospects
# Co-first authors
Received date: 2016-03-12
Accepted date: 2016-10-16
Online published: 2017-04-05
木灵藓科(Orthotrichaceae)是藓类植物中的第3大科。该科不仅种类多, 生态类型特殊, 而且是世界公认的多样化程度高、分类难度大、系统关系复杂的类群。当代木灵藓科植物分类系统学研究主要集中在该科的地区志编写和专属分类修订。目前, 除了热带美洲、热带非洲的变齿藓属(Zygodon)和火藓属(Schlotheimia)部分类群外, 木灵藓科主要类群的分类修订工作已基本完成, 但是有关亚科和属的划分和地位以及各属之间的关系等方面仍存在众多争议。木灵藓科分支系统学研究也不够系统全面, 有的仅应用了单个基因片段, 或者只涉及少数类群。因此, 需要基于更多的分子和形态学性状, 进一步开展世界木灵藓科植物的系统发育研究, 建立一个更趋自然的木灵藓科分类系统。
詹玲, 于晶, 郭水良, 曹同 . 木灵藓科分类与系统学研究:历史、问题和展望[J]. 植物学报, 2017 , 52(2) : 241 -253 . DOI: 10.11983/CBB16044
Orthotrichaceae is the third largest moss family according to species number. The family is rich in species and especially ecotypes but is also a taxon with high diversity, many taxonomic problems, and complex systematic relationship within its groups. The contemporary taxonomy and systematics of Orthotrichaceae has mainly focused on its local flora and monographs of relevant genera. The major genera have been taxonomically revised, except for some species names of Zygodon and Schlotheimia in tropical America and Africa. However, there are many disputes about subfamily divisions, systematic level of some genera, and relationships among different groups within the family. Moreover, previous relevant cladistics studies of Orthotrichaceae included only single gene fragments or did not include all taxa within the family. Therefore, more molecular and morphological data are needed for further phylogenetic research of the Orthotrichaceae family.
[1] | 陈邦杰 (1978). 中国藓类植物属志(下册). 北京: 科学出版社. pp. 1-331. |
[2] | 季必金 (2008). 贵州木灵藓科(Orthotrichaceae)植物分类及区系研究. 硕士论文. 贵阳: 贵州大学. pp. 16-42. |
[3] | 贾渝 (2012). 木灵藓科Orthotrichaceae. 见: 傅立国, 陈谭清, 郞楷永等. 中国高等植物, 第1卷. 青岛: 青岛出版社. pp. 10-252. |
[4] | 贾渝, 王庆华, 于宁宁 (2011). 木灵藓科Orthotrichaceae. 见: 吴鹏程, 贾渝. 中国苔藓志, 第5卷. 北京: 科学出版社. pp. 20-108. |
[5] | 王庆华 (2010). 卷叶藓属(Ulota)的分类修订. 博士论文. 北京: 中国科学院研究生院. pp. 17-149. |
[6] | 于宁宁 (2008). 裸帽藓属(Groutiella Steere)的分类学修订. 硕士论文. 济南: 山东师范大学. pp. 24-82. |
[7] | Allen B (1998). Five new species ofMacromitrium (Musci: Orthotrichaceae), with a key to the species of Macromitrium in Central America. Novon 8, 113-123. |
[8] | Allen B (2002). Moss Flora of Central America, Part 2. Encalyptaceae—Orthotrichaceae. St. Louis: Missouri Botanical Garden. pp. 1-699. |
[9] | Arnott GAW (1825). Disposition Méthodique des Espèces de Mousses.J Tastu 72, 4. |
[10] | Atwood JJ (2009). A taxonomic revision ofSchlotheimia subg. Stegotheca(Orthotrichaceae). Bryologist 112, 290-300. |
[11] | Bridel SE (1819). Muscologia recentiorum, seu analysis, historia, et description methodica omnium muscorum frondosorum hucusque cognitorum ad normam Hedwiii. S4, i-xviii, 1-120. |
[12] | Brotherus VF (1891). Contributions à la flore bryologique du Brésil.Acta Societatis Scientiarum Fennicae 19, 30. |
[13] | Brotherus VF (1909). Musci. In: Engler A, Prantl K, eds. Die Natürlichen Pflazenfamilien. Leipzig: W.Engelmann . pp. 277-1246. |
[14] | Brotherus VF (1925). Musci (Laubmoose) 2. In: Engler A, Prantl K, eds. Die Natürlichen Pflanzenfamilien Nebst ihren Gattungen und Wichtigeren Arten Insbesondere den Nutzpflanzen. Leipzig: W.Engelmann . pp. 1-542. |
[15] | Bruch P, Schimper WP (1838). Bryologia Europaea seu genera muscorum Europaeorum monographiche illustrate. Biodiversity Heritage Library. pp. 1864-1866. |
[16] | Buck WR, Goffinet B (2000). Morphology and classification of mosses. In: Shaw AJ, Goffinet B, eds. Bryophyte Biology. Cambridge: Cambridge University Press. pp. 71-123. |
[17] | Calabrese GM (2006). A taxonomic revision ofZygodon(Orthotrichaceae) in southern South America. Bryologist 109, 453-509. |
[18] | Calabrese GM, Elías MJ, Lara F (2006). A new species ofZygodon(Orthotrichaceae) from southern South America. J Bryol 28, 97-103. |
[19] | Calabrese GM, Muoz J (2008). Zygodon (Orthotrichaceae) in the Iberian Peninsula. Bryologist 111, 231-247. |
[20] | Cardot J (1908). La Flore Bryologique des Terres Magellaniques, de la Geogie du Sud et de I’Antarctide. In: Nordenskjöld O, ed. Wissenschaftliche Ergebnisse der Schwedischen Südpolar-expedition. Stockholm: MBLW- HOI Library. pp. 1-111. |
[21] | Churchill SP, Linares CE (1995). Prodromus bryologiae Novo-Granatensis: introducción a la flora de musgos de Colombia. Parte 2: Grimmiaceae a Trachypodaceae.Biblioteca José Jerónimo Triana 12, 455-924. |
[22] | Crosby MR, Magill RE (1981). A dictionary of mosses, third printing.Ann Mo Bot Gard 3, 43. |
[23] | Crosby MR, Magill RE, Allen B, He S (1999). A Checklist of the Mosses. Ann Mo Bot Gard. St. Louis: Missouri Botanical Garden. pp. 320. |
[24] | Crum H (1987). New species ofSphagnum from South America. J Hattori Bot Lab 63, 77-97. |
[25] | Crum HA, Anderson LE (1981). Mosses of Eastern North America, Vol. 1. New York: Columbia University Press. pp. 1-665. |
[26] | Damsholt K, Holmen K, Warncke E (1969). A list of the bryophytes of Denmark.Bot Tidsskr 65, 163-183. |
[27] | Dusén P (1905). Musci nonnulli novi e Fuegia et Patagonia reportati.Botaniska Notiser S, 299-310. |
[28] | Eddy A (1996). A Handbook of Malesian Mosses, Vol.3 (Splachnobryaceae to Leptostomataceae). London: Natural History Museum. pp. 1-277. |
[29] | Farge CL, Mishler BD, Wheeler JA, Wall DP, Johannes K (2000). Phylogenetic relationships within the haplolepidous mosses.Bryologist 103, 257-276. |
[30] | Fedosov VE, Ignatova EA (2010). Orthotrichum dagestanicum sp. nov. (Orthotrichaceae, Musci)—a new species from Dagestan (Eastern Caucasus).Arctoa 19, 69-74. |
[31] | Flora of North America Editorial Committee (2014). Bryophyta, part 2. In: Flora of North America. Cambridge: Cambridge University Press. Vol. 28, i- xxii, pp. 1-702. |
[32] | Gangulee HC (1976). Mosses of Eastern India and Adjacent Regions. Calcutta: Books and Allied Limited. pp. 1135-1462. |
[33] | Garilleti R, Lara F, Mazimpaka V (2006). Orthotrichum anodon (Orthotrichaceae), a new species from California, and its relationships to other Orthotricha with puckered capsule mouths.Bryologist 109, 188-196. |
[34] | Garilleti R, Mazimpaka V, Lara F (2012). NewUlota species with multicellular spores from southern South America. Bryologist 115, 585-600. |
[35] | Garilleti R, Shevock JR, Norris DH, Lara F (2011). Orthotrichum mazimpakanum sp. nov. and O. anodon(Orthotrichaceae), two similar species from California. Bryologist 114, 346-355. |
[36] | Goffinet B, Bayer RJ, Vitt DH (1998). Circumscription and phylogeny of the Orthotrichales (bryopsida) inferred fromrbcL sequence analyses. Am J Bot 85, 1324-1337. |
[37] | Goffinet B, Buck WR, Shaw AJ (2008). Morphology, anatomy, and classification of the Bryophyta. In: Goffinet B, Shaw AJ, eds. Bryophte Biology (2nd Ed.). Cambridge: Cambridge University Press. pp. 55-138. |
[38] | Goffinet B, Shaw AJ, Anderson LE, Mishler BD (1999). Peristome development in mosses in relation to systematics and evolution. V. Diplolepideae: Orthotrichaceae.Bryologist 102, 581-594. |
[39] | Goffinet B, Shaw AJ, Cox CJ, Wickett NJ, Boles S (2004). Phylogenetic inferences in the Orthotrichoideae (Orthotri- chaceae, Bryophyta) based on variation in four loci from all genomes.Ann Mo Bot Gard 98, 270-289. |
[40] | Goffinet B, Vitt DH (1998). Revised generic classification of the Orthotrichaceae based on a molecular phylogeny and comparative morphology. In: Bates JW, Ashton NW, Ashton NW, Duckett JG, eds. Bryology of the Twenty First Century. Leeds: Maney and British Bryological Society. pp. 143-159. |
[41] | Gradstein SR, Churchill SP, Salazar-Allen N (2001). Guide to the bryophytes of tropical America. Mem N Y Bot Gard 86, 1-577. |
[42] | Griffin DI (1990). Two new pentastichous species ofZygodon from high elevation in Venezuela. Cryptogamie Bryol 11, 163-168. |
[43] | Grout AJ (1935). Moss Flora of North America, North of Mexico, Vol. 2. Newfane, Vermont. pp. 106. |
[44] | Grout AJ (1944). Preliminary synopsis North American Macromitriae.Bryologist 47, 1-22. |
[45] | Guo SL, Cao T, Tan BC (2007a). Three new species records of Orthotrichaceae (Bryopsida) in China, with com- ments on their type specimens.Cryptogamie Bryol 28, 149-158. |
[46] | Guo SL, Cao T, Tan BC, Song GY (2007b). Taxonomic notes on Asian species of Orthotrichaceae (Bryopsida): Macromitrium with gymnostomous capsules. Gard B Sing- apore 58, 155-178. |
[47] | Guo SL, Enroth J, Koponen T (2007c). Bryophyte flora of Hunan Province, China. 11. Orthotrichaceae (Musci).Ann Bot Fennici 44, 1-34. |
[48] | Guo SL, Enroth J, Virtanen V (2004). Bryophyte flora of Hunan province, China. 10.Ulota gymnostoma sp. nova (orthotrichaceae). Ann Bot Fennici 41, 459-463. |
[49] | Guo SL, He S (2008a).Macromtrium hainanense, a new species of Orthotrichaceae from China. Bryologist 111, 505-509. |
[50] | Guo SL, He S (2008b).Macromitirum cavaleriei, a little known moss from China with four new synonyms. J Bryol 30, 264-270. |
[51] | Guo SL, He S (2014). Towards a new understanding ofMacromitrium nepalense(Orthotrichaceae), with two new synonyms. Bryologist 117, 15-21. |
[52] | Guo SL, Ma YH, Cao T, Lou YX (2012). A synopsis ofMacromitrium(Orthotrichaceae) in China. Cryptogamie Bryol 33, 341-355. |
[53] | Guo SL, Tan BC, Virtanen V (2006). Taxonomic and mor- phometric comments onMacromitrium blumei Nees ex Schwägrichen, M. zollingeri Mitten ex Bosch & Sande Lacoste and M. annamense Brotherus & Paris (Musci: Orthotrichaceae). Nova Hedwigia 82, 467-482. |
[54] | Hagen I (1908). Forarbejder til en norsk lovmosflora. I. Orthotrichceae.Det Kongelige Norske Videnskabers Selskabs Skrifter 13, 1-100. |
[55] | Hedwig J (1801). Species muscorum frondosorum descriptae et tabulis aeneis 77 coloratis illustratae. S1, i-xvi, 1-196. |
[56] | Heyn CC, Herrnstadt I (2004). The bryophyte flora of Israel and adjacent regions.Bryologist 3, 479. |
[57] | Hooker WJ (1824). Sketch of the characters of the species of Mosses.Edinburgh J Sci 1, 110-133. |
[58] | Ignatov M, Lewinsky J (1994). Bryophytes of Altai Mountains. II. The genera Amphidium Schimp., Orthotrichum Hedw. and Zygodon Hook. amp; amp; Tayl.(Orthotrichaceae, Musci). Arctoa 3, 29-57. |
[59] | Ignatov MS, Afonina OM (1992). Checklist of mosses of the former USSR.Arctoa 1, 1-85. |
[60] | Jia Y, He S, Guo SL (2011). Orthotrichaceae. In: Wu PC, Crosby MR, eds. Moss Flora of China, (English version) Erpodiaceae—Climaciaceae, 5. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. pp. 22-116. |
[61] | Koponen T, Enroth J (1992). Notes on the genusSchloth- eimia(Orthortrichaceae, Musci) in China. Bryobrothera 1, 277-282. |
[62] | Lara F, Garilleti R, Mazimpaka V (2006). Orthotrichum galiciae (Orthotrichaceae, Bryopsida), a new species from Ethiopia.Nova Hedwigia 83, 199-206. |
[63] | Lara F, Garilleti R, Mazimpaka V (2007). A peculiar newOrthotrichum species (Orthotrichaceae, Bryopsida) from central Argentina. Bot J Linn Soc 155, 477-482. |
[64] | Lara F, Garilleti R, Mazimpaka V (2016). Orthotrichum karoo (Orthotrichaceae), a new species with hyalineaw- ned leaves from southwestern Africa. Bryologist 112, 194-201. |
[65] | Lara F, Mazimpaka V, Medina R, Caparros R, Garilleti R (2010). Northeastern Turkey, an unnoticed but very important area for the Orthotrichaceae (Bryophyta).Nova Hedwigia 138, 165-180. |
[66] | Lewinsky J (1976). On the systematic position ofAmphidium Schimp. Lindbergia 3, 227-231. |
[67] | Lewinsky J (1977). The genusOrthotrichum—morpho- logical studies and evolutionary remarks. J Hattori Bot Lab 43, 31-61. |
[68] | Lewinsky J (1984a).Orthotrichum Hedw. in South America 1. Introduction and taxonomic revision of taxa with immersed stomata. Lindbergia 10, 65-94. |
[69] | Lewinsky J (1984b). The genusOrthotrichum Hedw.(Mus- ci) in Australasia: a taxonomic revision. J Hattori Bot Lab 56, 369-460. |
[70] | Lewinsky J (1987). Orthotrichum (Orthotrichaceae) in South America 2. Taxonomic revision of taxa with superficial stomata.Mem N Y Bot Gard 45, 326-370. |
[71] | Lewinsky J (1990). Zogodon Hook. & Tayl. in Australasia: a taxonomic revision.J Hattori Bot Lab 56, 369-460. |
[72] | Lewinsky J (1992). The genusOrthotrichum Hedw.(Orth- otrichaceae, Musci) in Southeast Asia. A taxonomic revision. J Hattori Bot Lab 72, 1-88. |
[73] | Lewinsky J (1993). The genusOrthotrichum Hedw.(Musci) in Hawaii. Bryobrothera 2, 61-69. |
[74] | Lewinsky J (1994). The genusPleurorthotrichum. Lindbergia 19, 11-24. |
[75] | Lewinsky J (1998). Orthotrichaceae. In: Nyholm E, ed. Illustrated Flora of Nordic Mosses, fasc 4. Copenhagen and Lund: Nordic Bryological Society. pp. 373-405. |
[76] | Lewinsky J, Crosby MR (1996). Orthotricum tuberculatum (Orthotrichaceae), a new genus and species from Gui- zhou, China.Novon 6, 1-5. |
[77] | Lewinsky J, Frisvoll A (1981). The genusOrthotrichum in Svalbard. Lindbergia 7, 2-9. |
[78] | Lewinsky J, Hedenäs L (1998). A cladistic analysis of the moss genusOrthotrichum. Bryologist 101, 519-555. |
[79] | Lewinsky-Haapasaari J (1994). Miscellaneous notes onOrthotrichum 5. Orthotrichum vicarium Laz. Lindbergia 19, 37-39. |
[80] | Lewinsky-Haapasaari J (1995). Orthotrichum notabile Le- winsky-Haapasaari, a new moss species from Sichuan, China.Lindbergia 20, 102-105. |
[81] | Lewinsky-Haapasaari J (1999a).Orthotrichum laxum, a new species from the Qinghai province, China. Lindbergia 24, 29-32. |
[82] | Lewinsky-Haapasaari J (1999b).Orthotrichum vermiferum, a new species from the Qinghai province of China, toge- ther with comments on Orthomitrium schofieldii(Orthotri- chaceae). Lindbergia 24, 33-37. |
[83] | Lewinsky-Haapasaari J, Tan BC (1995). Orthotrichum hallii sull. & lesq. new to Asia.Harvard Pap Bot 1, 1-5. |
[84] | Magill RE, Rooy JV (1998a). Bryophyta. Part 1 Musci, Fascicle 3, Erpodiaceae—Hookeriaceae. In: Flora of Southern Africa. Pretoria: Botanical Research Institute. pp. 462-475. |
[85] | Magill RE, Rooy JV (1998b). Bryophyta. Part 3, Erpodi- aceae-Hookeriaceae. In: Flora of Southern Africa. Pretoria: Botanical Research Institute. pp. 445-662. |
[86] | Malta N (1927). Die Ulota-Arten Süd-Amerikas.Acta Horti Botanici Universitatis Latviensis 2, 165-206. |
[87] | Marková I, Plášek V (2013). Zygodon dentatus (Limpr.) Kartt., a new moss species in the Bohemian Switzerland National Park (NW Bohemia, Czech Republic).Casopis Slezskeho Zemskeho Muzea 61, 251-254. |
[88] | Matcham HW, O'Shea BJ (2005). A review of the genusCodonoblepharon Schwagr.(Bryopsida: Orthotrichaceae). J Bryol 27, 129-135. |
[89] | Medina R, Lara F, Mazimpaka V, Garilleti R (2008). Orthotrichum norrisii (orthotrichaceae), a new epiphytic californian moss.Bryologist 111, 670-675. |
[90] | Medina R, Lara F, Mazimpaka V, Shevock JR, Garilleti R (2011). Orthotrichum pilosissimum (Orthotrichaceae), a new moss from arid areas of Nevada with unique axillary hairs. Bryologist 114, 316-324. |
[91] | Mitten W (1859a). Descriptions of some new species Musci from New Zealand and other parts of the Southern Hemisphere, together with an enumeration of the species collected in Tasmania by William Archer, arranged upon the plant proposed in the Musci Indiae Orientalis.J Proc Linn Soc Bot 4, 64-100. |
[92] | Mitten W (1859b). Musci Indiae Orientalis, an Enumeration of the Mosses of the East Indies.J Linn Soc Bot S1, 1-96. |
[93] | Mitten W (1869). Musci Austro-Americani.J Linn Soc Bot 12, 659. |
[94] | Mu?ller C (1849). Synopsis Muscorum Frondosorum Omnium Hucusque Cognitorum. Berolini: Sumptibus Alb. Foerstner. pp. 666-766. |
[95] | Mu?ller C (1878-1879). Musci fendleriani venezuelenses.Linnaea 42, 461-502. |
[96] | Mu?ller C (1896). Bryologia provinciae Schen-si Sinensis.Nuovo Giorn Bot Ital 3, 89-129. |
[97] | Mu?ller C (1897). Prodromus bryologiae Bolivianae.Nuovo Giorn Bot Ital 4, 113-172. |
[98] | Mu?ller C (1898). Symbolae ad bryologiam Australiae II.Hedwigia 37, 76-171. |
[99] | Müller C (1899). Contributiones ad Bryologiam austro- afram.Hedwigia 38, 52-155. |
[100] | Noguchi A (1968). Musci japonici. VIII. The genusOrthotri- chum. J Hattori Bot Lab 31, 115-129. |
[101] | Noguchi A, Iwatsuki Z (1989). Illustrated Moss Flora of Japan. Hattori Bot Lab. pp. 576-621. |
[102] | Norris DH, Robinson H (1981). Stoneobryum, a new genus of Orthotrichaceae from South Africa and southern Queensland.Bryologist 84, 95-99. |
[103] | Norris DH, Shevock JR, Goffinet B (2004). Orthotrichum kellmanii (Bryopsida, Orthotrichaceae), a remarkable new species from the central coast of California. Bryologist 107, 209-214. |
[104] | Nyholm E, Crum H (1956). Illustrated Moss Flora of Fennoscandia. Stockholm:Swedish Natural Science Research Council. pp. 309-313. |
[105] | Ochyra R, Smith RI, Bednarek-Ochrya H (2008). The illustrated Moss Flora of Antarctica.Bot J Linn Soc 164, 110-111. |
[106] | Othiokoba TH (2001). A peculiar newOrthotrichum species (Orthotrichaceae, Bryopsida) from central Argentina. Arctoa 10, 155-156. |
[107] | Plášek V, Dawicki J, Číhal L (2014). Orthotrichum pamiricum (Bryophyta), a new epiphytic moss species from Pamir Mountains in Central Asia.Turk J Bot 38, 754-762. |
[108] | Plášek V, Sawicki J (2012). Orthotrichum affine var.bohemicum-features and distribution. Nowellia Bryologica 44, 37-42. |
[109] | Plášek V, Sawicki J, Marková I, Wierzcholska S (2011). Orthotrichum affine var.bohemicum(Orthotrichaceae), a new variety of epiphytic moss from the Czech Republic. Acta Soc Bot Pol 80, 335-340. |
[110] | Plášek V, Sawicki J, Ochyra R (2015). New taxonomical arrangement of the traditionally conceived generaOrtho- trichum and Ulota(Orthotrichaceae, Bryophyta). Acta Mus Siles Sci Natur 64, 169-174. |
[111] | Plášek V, Sawicki J, TrávnÍčková V, Pasečná M (2009). Orthotrichum moravicum (Orthotrichaceae), a new moss from the Czech Republic. Bryologist 112, 329-336. |
[112] | Pluijm A (2001). Orthotrichum acuminatum H. philib., a Mediterranean moss new to the Netherlands.Lindbergia 26, 111-114. |
[113] | Ramsay HP, Vitt DH, Lewinsky-Haapasaari J (2006). Orthotrichaceae. In: Flora of Australia, Vol. 51. Mosses 1. Melbourne: ABRS, Canberra and CSIRO. pp. 187-244. |
[114] | Renauld F (1915). Histoire naturelle des plantes mousses viii Paris. In: Histoire Physique, Naturelle Et Politique De Madagascar. Paris: Librairie Hachette Et C1E. pp. 563. |
[115] | Robinson H (1964). New taxa and new records of bryophytes from Mexico and Central America.Bryologist 67, 446-458. |
[116] | Rooy JV, Wyk AEV (1992). A conspectus of the subfamily Macromitrioideae (Bryopsida: Orthotrichaceae) in Southern Africa. Bryologist 95, 205-215. |
[117] | Sainsbury GO (1955). A handbook of the New Zealand mosses.B Roy Soc New Zeal 5, 1-490. |
[118] | Sawicki J, Plášek V, Szczecińska M (2009). Preliminary studies on the phylogeny of the genusOrthotrichum inferred from nuclear ITS sequences. Ann Bot Fennici 46, 507-515. |
[119] | Sawicki J, Plášek V, Szczecińska M (2010). Molecular studies resolveNyholmiella(Orthotrichaceae) as a separate genus. J Syst Evol 48, 183-194. |
[120] | Sawicki J, Plášek V, Szczecińska M (2012). Molecular data do not support the current division ofOrthotrichum(Bryophyta) species with immersed stomata. J Syst Evol 50, 12-24. |
[121] | Schimper WP (1856). Corollarium Bryologiae Europaeae- Conspectum Diagnosticum Familiarum, Generum Et Spe- cierum, Adnotationes Novas Atque Emendationes. |
[122] | Smith AJ (2004). The Moss Flora of Britain & Ireland, 2nd edn. New York: Cambridge University Press. pp. 41-945. |
[123] | Tan BC, Jia Y (1997). Mosses of Qinghai-Tibetan plateau, China.J Hattori Bot Lab 82, 305-320. |
[124] | Vitt DH (1972). A monograph of the genusDrummondia. Can J Bot 50, 1191-1208. |
[125] | Vitt DH (1973). A revision of the genusOrthotrichum in North America, North of Mexico. Bryol Bibl 1, 1-208. |
[126] | Vitt DH (1976). A monograph of the genusMuelleriella Dusén. J Hattori Bot Lab 40, 91-113. |
[127] | Vitt DH (1979a). New taxa and new combinations in the Orthotrichaceae of Mexico.Bryologist 82, 1-19. |
[128] | Vitt DH (1979b). The status ofMuelleriella nom.cons., with Muelleriella atrata(Mitten) Vitt, an earlier name for M. engelii Vitt from Kerguelen Island.Lindbergia 5, 93-95. |
[129] | Vitt DH (1980a). The genusMacrocoma. I. Typification of names and taxonomy of the species.Bryologist 83, 405-436. |
[130] | Vitt DH (1980b). The genusMacrocoma. II. Geographical variation in the Macrocoma tenue-M. sullivantii species complex. Bryologist 83, 437-450. |
[131] | Vitt DH (1981). The generaLeiomitrium and Cardotiella gen. nova(Orthotrichaceae). J Hattori Bot Lab 49, 92-113. |
[132] | Vitt DH (1983). The New Zealand species of the pantropical genusMacromitrium (Orthotrichaceae: Musci): taxonomy, phylogeny and phytogeography. J Hattori Bot Lab 53, 1-94. |
[133] | Vitt DH (1989). The genusSchlotheimia(Orthotrichaceae: Bryopsida) in Australia and New Zealand. Bryologist 92, 282-298. |
[134] | Vitt DH (1990). Desmotheca (Orthotrichaceae): Gonwanan fragmentation and the origin of a Southeast Asian genus.Trop Bryol 3, 79-88. |
[135] | Vitt DH (1994). Orthotrichaceae. In: Sharp AJ, Crum H, Eckel ME, eds. The moss flora of Mexico. Mem N Y Bot Gard 69, 581-656. |
[136] | Vitt DH, Koponen T, Norris DH (1993). Bryophyte flora of the Huon Peninsula, Papua New Guinea. LIII.Ulota and Schlotheimia(Orthotrichaceae, Musci). Acta Bot Fennica 148, 5-25. |
[137] | Vitt DH, Koponen T, Norris DH (1995). Bryophyte flora of the Huon Peninsula, Papua New Guinea. LV.Desmoth- eca, Groutiella, Macrocoma and Macromitrium(Orthotrichaceae, Musci). Acta Bot Fennica 154, 1-94. |
[138] | Vitt DH, Ramsay HP (1985). TheMacromitrium complex in Australasia (Orthotrichaceae: Bryopsida). Part I. Taxonomy and phylogenetic relationships. J Hattori Bot Lab 59, 325-451. |
[139] | Wang QH, Jia Y (2012). A taxonomic revision of the Asian species ofUlota Mohr (Orthotrichaceae).Bryologist 115, 412-443. |
[140] | Wang QH, Jia Y (2014). Taxonomic status ofUlota tanganyikae P. de la Varde (Orthotrichaceae, Bryopsida) from Africa. J Bryol 36, 78-81. |
[141] | Wang QH, Jia Y (2016). A taxonomic revision ofUlota Mohr (Orthotrichaceae) in South and Central America. Plant Divers 38, 1-28. |
[142] | Wijk R Van Der, Margadant WD, Florschüetz PA (1967). Index Muscorum, Vol. IV. Utrecht: International Bureau for Plant Taxonomy and Nomenclature of the International Association for Plant Taxonomy. pp. 1-604. |
[143] | Wilbraham J (2008). New synonymy and typification in Afri- canSchlotheimia(Bryopsida: Orthotrichaceae). J Bryol 30, 196-200. |
[144] | Wilbraham J (2015). Annotated checklist and keys to the Orthotrichaceae of Malawi, together with new country records for East Africa. J Bryol 37, 87-95. |
[145] | Wilbraham J, Long DG (2005). Zygodon Hook. & Taylor andBryomaltaea Goffinet (Bryopsida: Orthotrichaceae) in the Sino-Himalaya. J Bryol 27, 329-342. |
[146] | Wilbraham J, Matcham HW (2010). Zygodon oeneus Herzog (Bryophyta: Orthotrichaceae) new to Uganda and Africa.J Bryol 32, 237-239. |
[147] | Wilbraham J, Price MJ (2013). A lectotype forMacromitrium cirrosum(Hedw.) Brid. J Bryol35, 119-122. |
[148] | Yu J, Guo SL, Ma YH, Cao T (2012). Taxonomic and mor- phometric comments onMacromitrium tosae Besch.(Or- thotrichaceae), with its four new synonyms.Bryologist 115, 388-401. |
[149] | Yu J, Guo SL, Ma YH, Cao T (2013). Macromitrium ousi- ense, a neglected Chinese moss species (Orthotrichaceae, Bryopsida) with new synonym and records.Nord J Bot 31, 339-343. |
[150] | Yu NN, Jia Y, Zhao JC (2011). Synonym and typifications inGroutiella tomentosa(Orthotrichaceae, Bryopsia). Novon 21, 290-293. |
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